This is a very good question.
I have my moon lights on a cycle that turns them off at 12:00 till 8:30 am giving the tank total darkness. This totally reduces the fish activity.
However in real life the shallow waters on a full moon lit night are not dark. I am sure that in the shallow waters it must be almost daylight. As fish are always on the hunt for food, I would think that non nocturnal fish would use a bright moon lit night to find an extra meal.
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